It is the year 2005. The battle between the Autobots and the Decepticons has escalated through their homeworld Cybertron, which the Decepticons have conquered. Without Optimus Prime after a conflict on Earth takes his life, the Autobots must face a destiny which they know nothing about. Megatron and a group of forsaken Decepticons have been reformed by the ultimate transformer, a planet consuming demon known as Unicron into even deadlier warriors. Galvatron, Scourge and Cyclonus must destroy the Autobot Matrix of Leadership for Unicron's glory or suffer the horrific destruction of Cybertron. However, Optimus Prime has decreed that an Autobot will rise from his rank and use the power of the Matrix to light the darkest hour of the Autobots. With Hot Rod facing responsibility for Prime's death, he feels he may be able to use the power of the Matrix to turn the tide of the Cybertronian Wars and stop Unicron. Until all are one, the future of the Autobots and the Decepticons is uncertain.